44. Umanzor, S., Meyer, A., Stamplis, Z., and Pryor, A. (2025). Effect of line spacing on blade phenotype and yields of farmed Alaria marginata from Alaska. Phycology 2025, 5, 89. https:// doi.org/10.3390/phycology5040089 

 

43. Dittrich, M.C., Meyer, L.A., Stekoll, M.S., Kelley, A., and Umanzor, S. (2005). Indoor cultivation protocols for Devaleraea mollis and Palmaria hecatensis from Alaska: effects of temperature, light, and nutrients. Journal of Applied Phycology. https://doi.org/10.1007/s10811-025-03594-x

 

42. Crimp, J., Meyer, L.A., Horstmann, L., Hollarsmith, J.A., Montiel, M., Lafarga-De La Cruz, F., Frederick, A.R., Licht, W, and Umanzor S. (2025). Effect of Macroalgal diet on growth rate and nutrition of the Pinto Abalone, Haliotis kamtschatkana. Journal of Shellfish Research, 44(1), 21-30. https://doi.org/10.2983/035.044.0103

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41. Jung, J. W., Dittrich, M., Kim, J. K., and Umanzor, S. (2025). Cost-effective nutrient supplementation strategies for land-based cultivation of Devaleraea mollis and Palmaria hecatensis. Journal of Applied Phycology. https://doi.org/10.1007/s10811-025-03480-6

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40. Almeida-Saá, A.C., Umanzor, S., Zertuche-González, J.A., Rangel-Mendoza, L.K., Ferreira-Arrieta, A., Cruz-López, R., and Sandoval-Gil, J.M. (2005). Comparative photophysiology and respiration of kelp gametophytes reveals species-specific thermo-tolerance to marine heatwaves. Mar Biol 172, 71. https://doi.org/10.1007/s00227-025-04630-7

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39. Jung, J.W., Dittrich, M., Umanzor, S., and Kim. J.K. (2025). Influence of Sargassum horneri extract (SHE) pre-treatment and nutrient availability on the thermal tolerance of Palmaria hecatensis. Journal of Applied Phycology. https://doi.org/10.1007/s10811-024-03419-3

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38. Umanzor, S., Jung, J.W., Dittrich, M., Kim, J.K., Tomco, P., Redman, Z.C., and Brandhuber, M. (2025). Divergent patterns of metabolite expression in red seaweeds Devaleraea mollis and Palmaria hecatensis following nitrate and ammonium supplementation. Life. https://doi.org/10.3390/life15020143

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37. Stephens, T. and Umanzor, S. (2024).  Comparative nutrients drawdown capacities of farmed kelps and implications of metabolic strategy and nutrient source. Journal of Phycology. 60(3):685-694. https://doi: 10.1111/jpy.13442

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36. Stephens, T., Li, Y., Yarish, C., Rogers, M.C., and Umanzor, S. (2024). Does Seawater Nitrogen Better Predict the Baseline Farmed Yield for Sugar Kelp (Saccharina latissima) Rather than the Final Yield? Phycology. 4(3):370-383. https://doi.org/10.3390/phycology4030020

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35. McArthur, M., Umanzor, S., Schram, J., and Konar, B. (2024). Environmental and biotic habitat attributes affect rocky intertidal community variability in glacially influenced estuaries. Journal of Experimental Marine Biology and Ecology. 58:152050. https://doi.org/10.1016/j.jembe.2024.152050

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34. Ham, G., Gabara, S., Chase, P., Konar, B., and Umanzor, S. (2024). Regional differences in the effects of watershed glacial coverage on the performance of Pacific Blue mussel, Mytilus trossulus. Journal of Experimental Marine Biology and Ecology. 580. https://doi.org/10.1016/j.jembe.2024.152043.

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33. Phelps, J., Dunton, K., Konar, B., Umanzor, S., Muth, A., and Iken, K. (2024). The effect of sedimentation on spore settlement and recruitment of the endemic Arctic kelp, Laminaria solidungula (Phaeophyceae). Journal of Phycology. http://doi.org/10.1111/jpy.13453

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32. Almeida-Saá, A.C., Umanzor, S., Zertuche-González, J.A., Cruz-López, R., Muñiz-Salazar, R., Ferreira-Arrieta, A., Bonet Melià, P., García-Pantoja, J.A. Rangel-Mendoza, L.K., Vivanco-Bercovich, M., Ruiz-Montoya, L., Guzmán-Calderón, and J.M. Sandoval-Gil, J.M. (2024). Bathymetric origin shapes the physiological responses of Pterygophora californica (Laminariales, Phaeophyceae) to deep marine heatwaves. Journal of Phycology. 00:1–20. https://doi.org/10.1111/jpy.13433

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31. Umanzor, S., Sandoval-Gil, J.M., and Conitz, J. (2023). Ecophysiological Responses of the Intertidal Seaweed Fucus distichus to temperature changes and reduced light driven by tides and glacial input. Estuaries and Coasts 46 (5): 1269 – 1279

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30. Huang, M., Robbins, K. R., Li, Y., Umanzor, S., Bailey, D., Aydlett, M., Schmutz, J., Grimwood, J., Yarish, C., Lindell, S., and Jannink, J. (2023). Genomic selection in algae with biphasic lifecycles: A Saccharina latissima (sugar kelp) case study. Frontiers in Marine Science, 10. https://doi.org/10.3389/fmars.2023.1040979

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29. Umanzor, S. and Stephens, T. (2023). Nitrogen and carbon removal capacity by farmed Alaria marginata and Saccharina latissima varies by species. Aquaculture Journal 3(1), 1-6; https://doi.org/10.3390/aquacj3010001

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28. Kite Powell, H., Ask, E., Augyte, S., Bailey, D., Decker, J., Goudey, C., Grebe, G., Li, Y., Lindell, S., Manganelli, D., Mary-Rivera, M., Ng, C., Roberson, L., Stekoll, M., Umanzor, S., and Yarish, C. (2023). Estimating production cost for large-scale seaweed farms. Applied Phycology. 3:1, 435-445 https://doi.org/10.1080/26388081.2022.2111271

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27. Han, S., Park, J.S., Umanzor, S., Yarish, C., and Kim, J.K. (2022). Effects of extraction methods for a new source of biostimulant from Sargassum horneri on the growth of economically important red algae, Neopyropia yezoensis. Scientific Reports 1–11

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26. Li, Y., Umanzor, S., Ng, C., Huang, M., Marty-Rivera, M., Bailey, D., Aydlett, M. et al. (2022). Skinny kelp (Saccharina angustissima) provides valuable genetics for the biomass improvement of farmed sugar kelp (Saccharina latissima). Journal of Applied Phycology 34, 2551–2563. https://doi.org/10.1007/s10811-022-02811-1

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25. Shelamoff, V., Umanzor, S., Layton, C., Tatsumi, M., Cameron, M.J., Wright, J.T., and Johnson, C.R. (2022). Ecosystem engineering kelp limits recruitment of mussels and microphytobenthic algae. Marine Biololy 1–14. https://doi.org/10.1007/s00227-022-04072-5

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24. Umanzor, S., Han, S., Song, H.-I., Park, J.S., Critchley, A.T., Yarish, C., and Jang, J.K. (2022). Ascertaining the interactions of brown seaweed-derived biostimulants and seawater temperature on spore release, germination, conchocelis, and newly formed blades of the commercially important red alga Neopyropia yezoensis? Algal Research. 64(3):102692

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23. Hossain, A., Apul, O. G., Dev, S., Das, S., Islam, S., Lai, C.-Y., Huntington, H., Kirisits, M. J., Umanzor, S., Chen, W.-T., Aggarwal, S., and Saleh, N.B. (2022). A symbiotic engineering approach for microplastic remediation in mariculture systems. ACS ES&T Engineering. https://doi.org/10.1021/acsestengg.1c00333

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22. Huang, M., Robbins, K.R., Li, Y., Umanzor, S., Marty-Rivera, M., Bailey, M., Yarish, C., Lindell, S., and Jannink, J-L. (2022). Simulation of sugar kelp (Saccharina latissima) breeding guided by practices to prioritize accelerated research gains. G3 Genes|Genomes|Genetics. https://doi.org/10.1093/g3journal/jkac003

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21. Umanzor, S., Han, S., Song H.I., Park, J.S., Critchley, A.T., Yarish, C., and Kim, J.K. (2022). Enhancements provided by the use of an Ascophyllum nodosum extract can be transferred through archeospores in the red alga Neopyropia yezoensis (Ueda) L.-E. Yang & J. Brodie. Aquatic Botany, 177: 103481

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20. Umanzor, S., Sandoval-Gil, J.M., Sanchez-Barredo, M., Ladah, L., Ramírez, M.M., and Zertuche-González, J.A (2021) Short-term stress responses and recovery of giant kelp (Macrocystis pyrifera, Laminariales, Phaeophyceae) juvenile sporophytes to a simulated marine heatwave and nitrate scarcity. Journal of Phycology, 57(5): 1604-1618. https://doi.org/10.1111/jpy.13189

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19. Augyte, S., Jannink, J-L., Mao, X., Huang, M., Robbins, K., Hare, M., Umanzor, S., Marty-Rivera, M. Li, Y., Lindell, S., Bailey, D., and Yarish, C. (2021). Kelp, Saccharina spp, population genetics in New England, US, for guiding a breeding program of thermally resilient strains. Bulletin of Japan Fisheries Research and Education Agency (50), 135-139

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18. Umanzor, S., Li, Y., Bailey, D., Augyte, S., Marty-Rivera, M., Huang, M., Jannink, J-L, Yarish, C., and Lindell, S. (2020). Comparative analysis of morphometric traits of farmed sugar kelp and skinny kelp (Saccharina spp.)  strains from the Northwest Atlantic. Journal of World Aquaculture Society 2021: 1–10. https://doi.org/10.1111/jwas.12783

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17. Umanzor, S., Li, Y., and Yarish, C. (2020). Effect of direct “seeding” binders and embryonic sporophyte sizes on the development of the sugar kelp, Saccharina latissima. Journal of Applied Phycology. https://doi.org/10.1007/s10811-020-02277-z

16. Mao, X., Augyte, S., Huang, M., Hare, M. P., Bailey, D., Umanzor, S., Marty-Rivera, M., Robbins, K. R., Yarish, C., Lindell, S., and Jannink, J.-L. (2020). Population genetics of sugar kelp in the Northwest Atlantic region using genome-wide markers. Frontiers in Marine Science. https://doi.org/10.3389/fmars.2020.00694

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15. Umanzor, S., Jang, S., Antosca, R. Critchley, A.T., Yarish, C., and Kim, J.K. (2020). Optimizing the application of selected biostimulants to enhance the growth of Eucheumatopsis isiformis, a carrageenophyte with commercial value, as grown in land-based nursery systems. Journal of Applied Phycology, 32, 1917–1922. https://doi.org/10.1007/s10811-020-02091-7

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14. Cabrera R., Díaz-Larrea J., Umanzor S., Clero L., Alfonso, Y., and Núñez-García L.G. (2020) Comparative growth and demographics of Thalassia testudinum meadows in Cuba using direct and reconstructive methods approaches to inform conservation efforts. International Journal of Recent Scientific Research 11:37446–37452. doi: 10.24327/IJRSR

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13. Augyte, S., G. Wikfors, S. Pitchford, M. Marty-Rivera, Umanzor, S., Yarish, C., Bailey, D., and Lindell, S. (2020). The application of flow cytometry for kelp meiospore isolation. Algal Research, 48: 101810, https://doi.org/10.1016/j.algal.2020.101810

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12. Umanzor, S., Shin, S. Marty-Rivera, M., Augyte, S., Yarish, C., and Kim, J.K. (2019) Exploratory evaluation of the effects of Kelpak® seaweed extract on cultivated kelp Saccharina spp. exposed to sublethal and lethal temperatures. Journal of World Aquaculture Society, 1-10. https://doi.org/10.1111/jwas.12687

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11. Umanzor, S., Ramírez, M.M., Sandoval-Gil, J.M., Zertuche-González, J.A, and Yarish, C. (2019). Photoacclimation and Photoprotection of Juvenile Sporophytes of Macrocystis pyrifera (Laminariales, Phaeophyceae) Under High-Light Conditions During Short-term Shallow-water Cultivation. Journal of Phycology, doi: 10.1111/jpy.12951

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10. Cabrera, R., Díaz-Larrea, J., Umanzor, S., and Núñez García, LG (2019). Using a macroalgal functional form approach to assess the level of disturbance of seagrass meadows in Bahía of Nuevitas, Cuba (2000 -2002). American Journal of Plant Sciences 10(11):2020-2033

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9. Cabrera, R., Umanzor, S., Díaz-Larrea, J., Araújo, P.G. (2019). Kappaphycus alvarezii (Rhodophyta): New record of an exotic species for the Caribbean Coast of Costa Rica. American Journal of Plant Sciences. https://doi.org/ 10.4236/ajps.2019.1010133

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8. Cabrera, R., Díaz-Larrea, J., and Umanzor, S. (2019). New records of marine macroalgae on the Caribbean Coast of Costa Rica. American Journal of Plant Sciences. https://doi.org/10.4236/ajps.2019.101012

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7. Umanzor, S., Shin, S., Marty-Rivera, M., Augyte, S., Yarish, C., and Kim, J.K. (2019). Preliminary assessment on the effects of the commercial seaweed extract, AMPEP, on growth and thermal tolerance of the kelp Saccharina spp. from the Northwest Atlantic. Journal of Applied Phycology. https://doi.org/10.1007/s10811-019-01852-3

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6. Umanzor, S., Ladah, L., Calderón-Aguilera, L.E., and Zertuche-González, J.A. (2019). Testing the relative importance of intertidal macroalgae as ecosystem engineers across extreme scenarios. Journal of Experimental Marine Biology and Ecology 511: 100–107. https://doi.org/10.3354/meps12355

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.5 Umanzor, S., Ladah, L., and Zertuche-González, J.A. (2018). Intertidal seaweeds modulate a contrasting response in understory macroalgal and microphytobenthic recruitment. Frontiers in Marine Science 5:296 .https://doi.org/10.3389/fmars.2018.00296

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4. Umanzor, S., Ladah, L., and Zertuche-González, J.A. (2017). The influence of species, density, and diversity of macroalgal aggregations on microphytobenthic settlement. Journal of Phycology 53 (5). https://doi.org/10.1111/jpy.12565

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3. Umanzor, S., Ladah, L., Calderón-Aguilera, L.E., and Zertuche-González, J.A. (2017). Intertidal macroalgae influence macroinvertebrate distribution across stress scenarios. Marine Ecology Progress Series 584: 67-77, https://doi.org/10.3354/meps12355

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2. Radulovich, R., Umanzor, S., Cabrera, R., and Mata, R. (2015). Tropical seaweeds for human food, their cultivation and its effect on biodiversity enrichment. Aquaculture 436: 40–46. https://doi.org/10.1016/j.aquaculture.2014.10.032

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1. Cheney, K.L., Cortesi, F., How, M.J., Wilson, N.G., Blomberg, S.P., Winters, A.E., Umanzor, S., and Marshall, N.J. (2014), Conspicuous visual signals do not coevolve with increased body size in marine sea slugs. Journal of Evolutionary Biology 27: 676–687. https://doi.org/10.1111/jeb.12348

 

Books and book chapters:

 

Radulovich, R. and Umanzor, S. (2020). Halophyte cultivation and use. In Handbook of Halophytes. From molecule to ecosystems towards biosaline agriculture. Grigore, M.N., Ed. Springer Nature, Switzerland. ISBN 978-3-030- 57635-6 (eBook)

 

Radulovich, R., Umanzor, S. and Cabrera, R. (2013). Algas Tropicales Cultivo y Uso como Alimento 2013 (Tropical Seaweeds: Cultivation and Use as Food) (1st ed.). Universidad de Costa Rica

 

Umanzor, S. and Conitz, J. (2025). Integrating Traditional Knowledge into Phyconomy of Temperate Regions. In Phyconomy of Temperate Regions. Critchley, A. Ed. Springer Nature (accepted)

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Selected non-peer-reviewed:

 

Hollarsmith, J., Umanzor, S., and Stephens, T. (2024). An overview of disease and pests on cultivated kelp, with a focus on Alaska. Proceedings of the 51st US-Japan Aquaculture Panel Symposium. NOAA Technical Memorandum NMFS-F/SPO-222

 

Umanzor, S., Good, M., Bobrycki, T., Kim, J.K., and Yarish, C. (2022). Building community capacity in site suitability assessments and determination of nutrient removal related to kelp mariculture. World Aquaculture Magazine. September. Vol. 53, number 3.

 

Umanzor, S., Shin, S.K., Augyte, S., Yarish, C., and Kim, J.K. (2018). Preliminary Experiments on Using the Seaweed Extract, AMPEP, to Enhance Thermal Tolerance and Growth of the Sugar Kelp, Saccharina latissima, from New England. Proceedings of the 46th US-Japan Aquaculture Panel Symposium. NOAA Technical Memorandum NMFS-F/SPO-222

 

Augyte S., Umanzor, S., Yarish C., and Lindell S. (2018). Enhancing marine ecosystem services via kelp aquaculture. ISAP Newsletter. December-2018 10–14

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Radulovich, R., Umanzor, S., Mata, R., and Elizondo, D. (2015). Aquatic Agriculture: Cultivating Floating Crops on Lakes. World Aquaculture Magazine, Mar: 62–67

 

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